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The Factors Of Organic Evolution
by [?]

[First published in The Nineteenth Century, for April and May, 1886.]

I.

Within the recollection of men now in middle life, opinion concerning the derivation of animals and plants was in a chaotic state. Among the unthinking there was tacit belief in creation by miracle, which formed an essential part of the creed of Christendom; and among the thinking there were two parties, each of which held an indefensible hypothesis. Immensely the larger of these parties, including nearly all whose scientific culture gave weight to their judgments, though not accepting literally the theologically-orthodox doctrine, made a compromise between that doctrine and the doctrines which geologists had established; while opposed to them were some, mostly having no authority in science, who held a doctrine which was heterodox both theologically and scientifically. Professor Huxley, in his lecture on “The Coming of Age of the Origin of Species,” remarks concerning the first of these parties as follows:–

“One-and-twenty years ago, in spite of the work commenced by Hutton and continued with rare skill and patience by Lyell, the dominant view of the past history of the earth was catastrophic. Great and sudden physical revolutions, wholesale creations and extinctions of living beings, were the ordinary machinery of the geological epic brought into fashion by the misapplied genius of Cuvier. It was gravely maintained and taught that the end of every geological epoch was signalised by a cataclysm, by which every living being on the globe was swept away, to be replaced by a brand-new creation when the world returned to quiescence. A scheme of nature which appeared to be modelled on the likeness of a succession of rubbers of whist, at the end of each of which the players upset the table and called for a new pack, did not seem to shock anybody.

I may be wrong, but I doubt if, at the present time, there is a single responsible representative of these opinions left. The progress of scientific geology has elevated the fundament principle of uniformitarianism, that the explanation of the past is to be sought in the study of the present, into the position of an axiom; and the wild speculations of the catastrophists, to which we all listened with respect a quarter of a century ago, would hardly find a single patient hearer at the present day.”

Of the party above referred to as not satisfied with this conception described by Professor Huxley, there were two classes. The great majority were admirers of the Vestiges of the Natural History of Creation–a work which, while it sought to show that organic evolution has taken place, contended that the cause of organic evolution, is “an impulse” supernaturally “imparted to the forms of life, advancing them, … through grades of organization.” Being nearly all very inadequately acquainted with the facts, those who accepted the view set forth in the Vestiges were ridiculed by the well-instructed for being satisfied with evidence, much of which was either invalid or easily cancelled by counter-evidence, and at the same time they exposed themselves to the ridicule of the more philosophical for being content with a supposed explanation which was in reality no explanation: the alleged “impulse” to advance giving us no more help in understanding the facts than does Nature’s alleged “abhorrence of a vacuum” help us to understand the ascent of water in a pump. The remnant, forming the second of these classes, was very small. While rejecting this mere verbal solution, which both Dr. Erasmus Darwin and Lamarck had shadowed forth in other language, there were some few who, rejecting also the hypothesis indicated by both Dr. Darwin and Lamarck, that the promptings of desires or wants produced growths of the parts subserving them, accepted the single vera causa assigned by these writers–the modification of structures resulting from modification of functions. They recognized as the sole process in organic development, the adaptation of parts and powers consequent on the effects of use and disuse–that continual moulding and re-moulding of organisms to suit their circumstances, which is brought about by direct converse with such circumstances.