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Transcendental Physiology
by
But our argument does not commit us to any such far-reaching speculation as this; which we introduce simply as suggested by it, not involved. All we are here concerned to show, is, that the deductive method aids us in interpreting some of the more general phenomena of development. That all homogeneous aggregates are in unstable equilibrium is a universal truth, from which is deducible the instability of every organic germ. From the known sensitiveness of organic compounds to chemical, thermal, and other disturbing forces, we further infer the unusual instability of every organic germ–a proneness far beyond that of other homogeneous aggregates to lapse into a heterogeneous state. By the same line of reasoning we are led to the additional inference, that the first divisions into which a germ resolves itself, being severally in a state of unstable equilibrium, are similarly prone to undergo further changes; and so on continuously. Moreover, we have found it to be equally an a priori conclusion, that as, in all other cases, the loss of homogeneity is due to the different degrees and kinds of force brought to bear on the different parts; so, in this case too, difference of circumstances is the primary cause of differentiation. Add to which, that as the several changes undergone by the respective parts thus diversely acted upon, are changes which do not destroy their vital activity, they must be changes which bring that vital activity into subordination to the incident forces–they must be adaptations; and the like must be in some sense true of all the subsequent changes. Thus by deductive reasoning we get some insight into the method of organization. However unable we are, and probably ever shall be, to comprehend the way in which a germ is made to take on the special form of its race, we may yet comprehend the general principles which regulate its first modifications; and, remembering the unity of plan so conspicuous throughout nature, we may suspect that these principles are in some way concerned in succeeding modifications.
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A controversy now going on among zoologists, opens yet another field for the application of the deductive method. We believe that the question whether there does or does not exist a necessary correlation among the several parts of an organism is determinable a priori.
Cuvier, who first asserted this necessary correlation, professed to base his restorations of extinct animals upon it. Geoffroy St. Hilaire and De Blainville, from different points of view, contested Cuvier’s hypothesis; and the discussion, which has much interest as bearing on paleontology, has been recently revived under a somewhat modified form: Professors Huxley and Owen being respectively the assailant and defender of the hypothesis.
Cuvier says–“Comparative anatomy possesses a principle whose just development is sufficient to dissipate all difficulties; it is that of the correlation of forms in organized beings, by means of which every kind of organized being might, strictly speaking, be recognized by a fragment of any of its parts. Every organized being constitutes a whole, a single and complete system, whose parts mutually correspond and concur by their reciprocal reaction to the same definite end. None of these parts can be changed without affecting the others; and consequently each taken separately, indicates and gives all the rest.” He then gives illustrations: arguing that the carnivorous form of tooth necessitating a certain action of the jaw, implies a particular form in its condyles; implies also limbs fit for seizing and holding prey; therefore implies claws, a certain structure of the leg-bones, a certain form of shoulder-blade. Summing up he says, that “the claw, the scapula, the condyle, the femur, and all the other bones, taken separately, will give the tooth or one another; and by commencing with any one, he who had a rational conception of the laws of the organic economy, could reconstruct the whole animal.”