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“By degrees, in the midst of the chaos of the rising society, small aggregations are formed which feel the want of alliance and union with each other…. Soon inequality of strength is displayed among neighbouring aggregations. The strong tend to subjugate the weak, and usurp at first the rights of taxation and military service. Thus political authority leaves the aggregations which first instituted it, to take a wider range.”

That is to say, the small tribes, clans, or feudal groups, sprung mostly from a common stock, and long held in contact as occupants of adjacent lands, gradually get united in other ways than by kinship and proximity.

A further series of changes begins now to take place, to which, as before, we find analogies in individual organisms. Returning to the Hydrozoa, we observe that in the simplest of the compound forms the connected individuals are alike in structure, and perform like functions; with the exception that here and there a bud, instead of developing into a stomach, mouth, and tentacles, becomes an egg-sac. But with the oceanic Hydrozoa this is by no means the case. In the Calycophoridae some of the polypes growing from the common germ, become developed and modified into large, long, sack-like bodies, which, by their rhythmical contractions, move through the water, dragging the community of polypes after them. In the Physophoridae a variety of organs similarly arise by transformation of the budding polypes; so that in creatures like the Physalia, commonly known as the “Portuguese Man-of-war,” instead of that tree-like group of similar individuals forming the original type, we have a complex mass of unlike parts fulfilling unlike duties. As an individual Hydra may be regarded as a group of Protozoa which have become partially metamorphosed into different organs; so a Physalia is, morphologically considered, a group of Hydrae of which the individuals have been variously transformed to fit them for various functions.

This differentiation upon differentiation is just what takes place during the evolution of a civilized society. We observed how, in the small communities first formed, there arises a simple political organization: there is a partial separation of classes having different duties. And now we have to observe how, in a nation formed by the fusion of such small communities, the several sections, at first alike in structures and modes of activity, grow unlike in both–gradually become mutually-dependent parts, diverse in their natures and functions.

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The doctrine of the progressive division of labour, to which we are here introduced, is familiar to all readers. And further, the analogy between the economical division of labour and the “physiological division of labour,” is so striking as long since to have drawn the attention of scientific naturalists: so striking, indeed, that the expression “physiological division of labour,” has been suggested by it. It is not needful, therefore, to treat this part of the subject in great detail. We shall content ourselves with noting a few general and significant facts, not manifest on a first inspection.

Throughout the whole animal kingdom, from the Coelenterata upwards, the first stage of evolution is the same. Equally in the germ of a polype and in the human ovum, the aggregated mass of cells out of which the creature is to arise, gives origin to a peripheral layer of cells, slightly differing from the rest which they include; and this layer subsequently divides into two–the inner, lying in contact with the included yelk, being called the mucous layer, and the outer, exposed to surrounding agencies, being called the serous layer: or, in the terms used by Prof. Huxley, in describing the development of the Hydrozoa–the endoderm and ectoderm. This primary division marks out a fundamental contrast of parts in the future organism. From the mucous layer, or endoderm, is developed the apparatus of nutrition; while from the serous layer, or ectoderm, is developed the apparatus of external action. Out of the one arise the organs by which food is prepared and absorbed, oxygen imbibed, and blood purified; while out of the other arise the nervous, muscular, and osseous systems, by the combined actions of which the movements of the body as a whole are effected. Though this is not a rigorously-correct distinction, seeing that some organs involve both of these primitive membranes, yet high authorities agree in stating it as a broad general distinction. Well, in the evolution of a society, we see a primary differentiation of analogous kind, which similarly underlies the whole future structure. As already pointed out, the only manifest contrast of parts in primitive societies, is that between the governing and the governed. In the least organized tribes, the council of chiefs may be a body of men distinguished simply by greater courage or experience. In more organized tribes, the chief-class is definitely separated from the lower class, and often regarded as different in nature–sometimes as god-descended. And later, we find these two becoming respectively freemen and slaves, or nobles and serfs. A glance at their respective functions, makes it obvious that the great divisions thus early formed, stand to each other in a relation similar to that in which the primary divisions of the embryo stand to each other. For, from its first appearance, the warrior-class, headed by chiefs, is that by which the external acts of the society are carried on: alike in war, in negotiation, and in migration. Afterwards, while this upper class grows distinct from the lower, and at the same time becomes more and more exclusively regulative and defensive in its functions, alike in the persons of kings and subordinate rulers, priests, and soldiers; the inferior class becomes more and more exclusively occupied in providing the necessaries of life for the community at large. From the soil, with which it comes in most direct contact, the mass of the people takes up, and prepares for use, the food and such rude articles of manufacture as are known; while the overlying mass of superior men, maintained by the working population, deals with circumstances external to the community–circumstances with which, by position, it is more immediately concerned. Ceasing by-and-by to have any knowledge of, or power over, the concerns of the society as a whole, the serf-class becomes devoted to the processes of alimentation; while the noble class, ceasing to take any part in the processes of alimentation, becomes devoted to the co-ordinated movements of the entire body-politic.