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You see, there was hardly ever a big storm on land that didn’t bring at least one or two new birds of some sort or other to the islands. Naturally, too, the newcomers landed always on the first shore they could sight; and so at the present day the greatest number of species is found on the two easternmost islands nearest the mainland, which have forty kinds of land-birds, while the central islands have but thirty-six, and the western only twenty-nine. It would have been quite different, of course, if the birds came mainly from America with the trade winds and the Gulf Stream, as I at first anticipated. In that case, there would have been most kinds in the westernmost islands, and fewest stragglers in the far eastern. But your own naturalists have rightly seen that the existing distribution necessarily implies the opposite explanation.

Birds, I early noticed, are always great carriers of fruit-seeds, because they eat the berries, but don’t digest the hard little stones within. It was in that way, I fancy, that the Portugal laurel first came to my islands, because it has an edible fruit with a very hard seed; and the same reason must account for the presence of the myrtle, with its small blue berry; the laurustinus with its currant-like fruit; the elder-tree, the canary laurel, the local sweet-gale, and the peculiar juniper. Before these shrubs were introduced thus unconsciously by our feathered guests, there were no fruits on which berry-eating birds could live; but now they are the only native trees or large bushes on the islands–I mean the only ones not directly planted by you mischief-making men, who have entirely spoilt my nice little experiment.

It was much the same with the history of some among the birds themselves. Not a few birds of prey, for example, were driven to my little archipelago by stress of weather in its very early days; but they all perished for want of sufficient small quarry to make a living out of. As soon, however, as the islands had got well stocked with robins, black-caps, wrens, and wagtails, of European types–as soon as the chaffinches had established themselves on the seaward plains, and the canary had learnt to nest without fear among the Portugal laurels–then buzzards, long-eared owls, and common barn-owls, driven westward by tempests, began to pick up a decent living on all the islands, and have ever since been permanent residents, to the immense terror and discomfort of our smaller song-birds. Thus the older the archipelago got the less chance was there of local variation taking place to any large degree, because the balance of life each day grew more closely to resemble that which each species had left behind it in its native European or African mainland.

I said a little while ago we had no mammal in the islands. In that I was not quite strictly correct. I ought to have said, no terrestrial mammal. A little Spanish bat got blown to us once by a rough nor’easter, and took up its abode at once among the caves of our archipelago, where it hawks to this day after our flies and beetles. This seemed to me to show very conspicuously the advantage which winged animals have in the matter of cosmopolitan dispersion; for while it was quite impossible for rats, mice, or squirrels to cross the intervening belt of three hundred leagues of sea, their little winged relation, the flitter-mouse, made the journey across quite safely on his own leathery vans, and with no greater difficulty than a swallow or a wood-pigeon.

The insects of my archipelago tell very much the same story as the birds and the plants. Here, too, winged species have stood at a great advantage. To be sure, the earliest butterflies and bees that arrived in the fern-clad period were starved for want of honey; but as soon as the valleys began to be thickly tangled with composites, harebells, and sweet-scented myrtle bushes, these nectar-eating insects established themselves successfully, and kept their breed true by occasional crosses with fresh arrivals blown to sea afterwards. The development of the beetles I watched with far greater interest, as they assumed fresh forms much more rapidly under their new conditions of restricted food and limited enemies. Many kinds I observed which came originally from Europe, sometimes in the larval state, sometimes in the egg, and sometimes flying as full-grown insects before the blast of the angry tempest. Several of these changed their features rapidly after their arrival in the islands, producing at first divergent varieties, and finally, by dint of selection, acting in various ways, through climate, food, or enemies, on these nascent forms, evolving into stable and well-adapted species. But I noticed three cases where bits of driftwood thrown up from South America on the western coasts contained the eggs or larvae of American beetles, while several others were driven ashore from the Canaries or Madeira; and in one instance even a small insect, belonging to a type now confined to Madagascar, found its way safely by sea to this remote spot, where, being a female with eggs, it succeeded in establishing a flourishing colony. I believe, however, that at the time of its arrival it still existed on the African continent, but becoming extinct there under stress of competition with higher forms, it now survives only in these two widely separated insular areas.