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Let us suppose that in a region now covered by wide ocean, there begins one of those great and gradual upheavals by which new continents are formed. To be precise, let us say that in the South Pacific, midway between New Zealand and Patagonia, the sea-bottom has been little by little thrust up toward the surface, and is about to emerge. What will be the successive phenomena, geological and biological, which are likely to occur before this emerging sea-bottom has become another Europe or Asia? In the first place, such portions of the incipient land as are raised to the level of the waves, will be rapidly denuded by them: their soft substance will be torn up by the breakers, carried away by the local currents, and deposited in neighbouring deeper water. Successive small upheavals will bring new and larger areas within reach of the waves; fresh portions will each time be removed from the surfaces previously denuded; and further, some of the newly-formed strata, being elevated nearly to the level of the water, will be washed away and re-deposited. In course of time the harder formations of the upraised sea-bottom will be uncovered. These, being less easily destroyed, will remain permanently above the surface; and at their margins will arise the usual breaking down of rocks into beach-sand and pebbles. While in the slow course of this elevation, going on at the rate of perhaps two or three feet in a century, most of the sedimentary deposits produced will be again and again destroyed and reformed; there will, in those adjacent areas of subsidence which accompany areas of elevation, be more or less continuous successions of sedimentary deposits lying on the pre-existing ocean bed. And now, what will be the character of these strata, old and new? They will contain scarcely any traces of life. The deposits that had previously been slowly formed at the bottom of this wide ocean, would be sprinkled with fossils of but few species. The oceanic Fauna is not a rich one; its hydrozoa do not admit of preservation; and the hard parts of its few kinds of molluscs and crustaceans and insects are mostly fragile. Hence, when the ocean-bed was here and there raised to the surface–when its strata of sediment with their contained organic fragments were torn up and long washed about by the breakers before being re-deposited–when the re-deposits were again and again subject to this violent abrading action by subsequent small elevations, as they would mostly be; what few fragile organic remains they contained, would be in nearly all cases destroyed. Thus such of the first-formed strata as survived the repeated changes of level, would be practically “azoic;” like the Cambrian of our geologists. When by the washing away of the soft deposits, the hard sub-strata had been exposed in the shape of rocky islets, and a footing had thus been furnished, the pioneers of a new life might be expected to make their appearance. What would they be? Not any of the surrounding oceanic species, for these are not fitted for a littoral life; but species flourishing on some of the far-distant shores of the Pacific. Of such, the first to establish themselves would be sea-weeds and zoophytes; because the most readily conveyed on floating wood, etc., and because when conveyed they would find fit food. It is true that Cirrhipeds and Lamellibranchs, subsisting on the minute creatures which everywhere people the sea, would also find fit food. But the chances of early colonization are in favour of species which, multiplying by agamogenesis, can people a whole shore from a single germ; and against species which, multiplying only by gamogenesis, must be introduced in considerable numbers that some may propagate. Thus we infer that the earliest traces of life left in the sedimentary deposits near these new shores, will be traces of life as humble as that indicated in the most ancient rocks of Great Britain and Ireland. Imagine now that the processes above indicated, continue–that the emerging lands become wider in extent, and fringed by higher and more varied shores; and that there still go on those ocean-currents which, at long intervals, convey from far distant shores immigrant forms of life. What will result? Lapse of time will of course favour the introduction of such new forms: admitting, as it must, of those combinations of fit conditions, which can occur only after long intervals. Moreover, the increasing area of the islands, individually and as a group, implies increasing length of coast, and therefore a longer line of contact with the streams and waves which bring drifting masses bearing germs of fresh life. And once more, the comparatively-varied shores, presenting physical conditions which change from mile to mile, will furnish suitable habitats for more numerous species. So that as the elevation proceeds, three causes conspire to introduce additional marine plants and animals. To what classes will the increasing Fauna be for a long period confined? Of course, to classes of which individuals, or their germs, are most liable to be carried far away from their native shores by floating sea-weed or drift-wood; to classes which are also least likely to perish in transit, or from change of climate; and to those which can best subsist around coasts comparatively bare of life. Evidently then, corals, annelids, inferior molluscs, and crustaceans of low grade, will chiefly constitute the early Fauna. The large predatory members of these classes, will be later in establishing themselves; both because the new shores must first become well peopled by the creatures they prey on, and because, being more complex, they, or their ova, must be less likely to survive the journey, and the change of conditions. We may infer, then, that the strata deposited next after the almost “azoic” strata, would contain the remains of invertebrata, allied to those found near the shores of Australia and South America. Of such invertebrate remains, the lower beds would furnish comparatively few genera, and those of relatively low types; while in the upper beds the number of genera would be greater, and the types higher: just as among the fossils of our Silurian system. As this great geologic change slowly advanced through its long history of earthquakes, volcanic disturbances, minor upheavals and subsidences–as the extent of the archipelago became greater and its smaller islands coalesced into larger ones, while its coast-line grew still longer and more varied, and the neighbouring sea more thickly inhabited by inferior forms of life; the lowest division of the vertebrata would begin to be represented. In order of time, fish would naturally come later than the lower invertebrata; both as being less likely to have their ova transported across the waste of waters, and as requiring for their subsistence a pre-existing Fauna of some development. They might be expected to make their appearance along with the predaceous crustaceans; as they do in the uppermost Silurian rocks. And here, too, let us remark, that as, during this long epoch we have been describing, the sea would have made great inroads on some of the newly-raised lands which had remained stationary; and would probably in some places have reached masses of igneous or metamorphic rocks; there might, in course of time, arise by the decomposition and denudation of such rocks, local deposits coloured with oxide of iron, like our Old Red Sandstone. And in these deposits might be buried the remains of the fish then peopling the neighbouring sea.